1. Department of Aquatic Animal Medicine, College of Fisheries, Huazhong Agricultural University, Wuhan 430070, People’s Republic of China.
2. Freshwater Aquaculture Collaborative Innovation Center of Hubei Province, Wuhan 430070, People’s Republic of China.
3. Key Laboratory of Freshwater Animal Breeding, Ministry of Agriculture, Wuhan 430070, People’s Republic of China.