1. Pathology Division, Nansei Field Station Fisheries Technology Institute, Japan Fisheries Research and Education Agency Minami‐Ise Mie Japan
2. Pathology Division, Tamaki Field Station Fisheries Technology Institute, Japan Fisheries Research and Education Agency Tamaki Mie Japan
3. National Reference Laboratory for Fish Diseases, Wageningen Bioveterinary Research Wageningen University & Research Lelystad The Netherlands