1. Saitama Prefectural Chuo Livestock Hygiene Service Center, 107–1 Besshocho, Kita-ku, Saitama, Saitama 331–0821, Japan
2. The United Graduate School of Veterinary Sciences, Gifu University, 1–1 Yanagido, Gifu, Gifu 501–1193, Japan
3. Department of Infectious Diseases, National Center for Global Health and Medicine, 1–21–1 Toyama, Shinjuku-ku, Tokyo 162–8655, Japan
4. Department of Animal and Food Hygiene, Obihiro University of Agriculture and Veterinary Medicine, 11 Nishi 2-sen, Inada-cho, Obihiro, Hokkaido 080–8555, Japan
5. Fukuoka-Chuo Prefectural Institute of Animal Health, 4–14–5 Hakozakifuto, Higashi-ku, Fukuoka, Fukuoka 812–0051, Japan
6. Graduate School of Life and Environmental Sciences, University of Tsukuba, 1–1–1 Tennodai, Tsukuba, Ibaraki 305–8572, Japan
7. Honey Bee Research Unit, Animal Breeding and Reproduction Research Division, NARO Institute of Livestock and Grassland Science, National Agriculture and Food Research Organization, 2 Ikenodai, Tsukuba, Ibaraki 305–0901, Japan
8. Bacterial and Parasitic Diseases Research Division, National Institute of Animal Health, National Agriculture and Food Research Organization, 3–1–5 Kannondai, Tsukuba, Ibaraki 305–0856, Japan