1. Laboratory of Veterinary Microbiology, Graduate School of Medicine and Veterinary Medicine, University of Miyazaki, 1-1 Gakuenkibanadai Nishi, Miyazaki, Miyazaki 889-2192, Japan
2. Laboratory of Veterinary Microbiology, Department of Veterinary Science, Faculty of Agriculture, University of Miyazaki, 1-1 Gakuenkibanadai Nishi, Miyazaki, Miyazaki 889-2192, Japan
3. Center for Animal Disease Control, University of Miyazaki, 1-1 Gakuenkibanadai Nishi, Miyazaki, Miyazaki 889-2192, Japan