1. Laboratory of Food Microbiology and Food Safety, School of Veterinary Medicine, Rakuno Gakuen University, Hokkaido, Japan
2. Biological Science Laboratories, Kao Corporation, Tochigi, Japan
3. Division of Bioresources, Hokkaido University International Institute for Zoonosis Control, Hokkaido, Japan
4. International Collaboration Unit, Hokkaido University International Institute for Zoonosis Control, Hokkaido, Japan
5. Institute for Vaccine Research and Development, Hokkaido University, Hokkaido, Japan