1. Aichi Prefectural Chuo Livestock Hygiene Service Center, 1-306 Jizono, Miaicho, Okazaki, Aichi 444-0805, Japan
2. Tochigi Prefectural Central Livestock Health and Hygiene Center, 6-8 Hiraide-kogyodanchi, Utsunomiya, Tochigi 321-0905, Japan
3. Division of Bacterial and Parasitic Disease, National Institute of Animal Health, National Agriculture and Food Research Organization, 3-1-5 Kannondai, Tsukuba, Ibaraki 305-0856, Japan
4. The United Graduate School of Veterinary Sciences, Gifu University, 1–1 Yanagido, Gifu, Gifu 501-1193, Japan
5. Division of Pathology and Pathophysiology, National Institute of Animal Health, National Agriculture and Food Research Organization, 3-1-5 Kannondai, Tsukuba, Ibaraki 305-0856, Japan
6. Department of Veterinary Science, Graduate School of Life and Environmental Sciences, Osaka Prefecture University, 1-58 Rinku-oraikita, Izumisano, Osaka 598-8531, Japan