1. Department of Disease Control, Faculty of Veterinary Medicine, Hokkaido University, Sapporo, Hokkaido 060-0818, Japan
2. Department of Advanced Pharmaceutics, Faculty of Veterinary Medicine, Hokkaido University, Sapporo, Hokkaido 060-0818, Japan
3. Bacterial and Parasitic Disease Research Division, National Institute of Animal Health, Tsukuba, Ibaraki 305-0856, Japan
4. Department of Antibody Drug Development, Tohoku University Graduate School of Medicine, 2-1 Seiryo-machi, Aoba-ku, Sendai, Miyagi 980-8575, Japan
5. New Industry Creation Hatchery Center, Tohoku University, 2-1 Seiryo-machi, Aoba-ku, Sendai, Miyagi 980-8575, Japan
6. Division of Bioresources, Research Center for Zoonosis Control, Hokkaido University, Sapporo, Hokkaido 001-0019, Japan