1. Cooperative Department of Veterinary Medicine, Iwate University, Morioka, Iwate 020–8550, Japan
2. Field Science Center, Faculty of Agriculture, Iwate University, Shizukuishi, Iwate 020–0581, Japan
3. Center for Biotechnology, Agriculture and Forestry University, Rampur, Chitwan, Nepal
4. Current affiliation: Department of Clinical Veterinary Medicine, Obihiro University of Agriculture and Veterinary Medicine, Inada, Obihiro, Hokkaido 080–8555, Japan