1. Center for Macroecology, Evolution and Climate, Natural History Museum of Denmark, University of Copenhagen, DK-2100 Copenhagen O, Denmark.
2. Section for Ecoinformatics and Biodiversity, Department of Bioscience, Aarhus University, DK-8000 Aarhus C, Denmark.
3. Center for Biodiversity Dynamics in a Changing World (BIOCHANGE), Department of Bioscience, Aarhus University, DK-8000 Aarhus C, Denmark.