1. Department of Forestry and Wildlife Management, Faculty of Applied Ecology and Agricultural Sciences, Hedmark University College, Campus Evenstad, NO-2480 Koppang, Norway
2. Finnish Food Safety Authority Evira, Production Animal and Wildlife Health Research Unit (FINPAR), Elektroniikkatie 3, FI-90590, Oulu, Finland
3. Faculty of Veterinary Medicine, University of Helsinki, PO Box 66, FI-00014 University of Helsinki, Helsinki, Finland