1. Animal Science and Genetics, Tasmanian Institute of Agriculture, School of Land and Food, Faculty of Science, Engineering and Technology, University of Tasmania, Private Bag 54, Hobart, Tasmania, 7001, Australia
2. Veterinary & Biomedical Sciences, College of Public Health, Medical and Veterinary Sciences, Division of Tropical Health and Medicine, James Cook University, Townsville, Queensland, 4811, Australia
3. Red Meat and Sheep Development Centre, NSW Department of Primary Industries, Cowra, 2794, Australia(B.W.B. Holman)