1. 1Department of Biotechnology, The University of Tokyo, Yayoi, Bunkyo-ku, Tokyo, Japan
2. 2NODAI Genome Research Center, Tokyo University of Agriculture, Setagaya-ku, Tokyo, Japan
3. 3Department of Bioscience, Tokyo University of Agriculture, Setagaya-ku, Tokyo, Japan
4. †Present address: Graduate School of Frontier Sciences, The University of Tokyo, 5-1-5 Kashiwanoha, Kashiwa, Chiba 277-8562, Japan.