1. 1College of William & Mary, Department of Biology, Integrated Science Center Rm 3035, 540 Landrum Dr Williamsburg, VA 23185, USA
2. 2Department of Ecology, Evolution, and Behavior, University of Minnesota-Twin Cities, 100 Ecology Building, 1987 Upper Buford Circle, Saint Paul, MN 55108, USA
3. 3BioTechnology Institute, University of Minnesota-Twin Cities, 140 Gortner Labs, 1479 Gortner Avenue, St Paul, MN 55108, USA
4. 4Department of Biology, New Mexico State University, Box 30001, MSC 3AF, Las Cruces, NM 88003, USA