1. Department of Botany and Plant Pathology, Oregon State University, Corvallis, OR, USA
2. USDA-ARS Forage Seed and Cereal Research Unit, Corvallis, OR, USA
3. Department of Crop and Soil Science, Oregon State University, Corvallis, OR, USA
4. Present address: Department of Biochemistry, Molecular Biology and Biophysics, University of Minnesota, St. Paul, MN, USA.