1. Laboratory of Microbiology, Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo 060-0818, Japan
2. Institute of Virology and Immunology IVI, Sensemattstrasse 293, CH-3147 Mittelhäusern, Switzerland
3. Research Center for Zoonosis Control, Hokkaido University, Sapporo 001-0020, Japan
4. Global Station for Zoonosis Control, Global Institution for Collaborative Research and Education (GI-CoRE), Hokkaido University, Sapporo 001-0020, Japan