1. Department of Veterinary Preclinical Science, University of Liverpool, Brownlow Hill and Crown Street, Liverpool L69 7ZJ, UK
2. Department of Veterinary Pathology, University of Liverpool, Leahurst, Neston, South Wirral CH64 7TE, UK
3. Department of Veterinary Clinical Science, University of Liverpool, Leahurst, Neston, South Wirral CH64 7TE, UK