1. Department of Biotechnology, Mokwon University, Doan-dong 800, Seo-gu, Daejeon 302-729, Republic of Korea
2. Institute of Microbial Ecology and Resources, Mokwon University, Doan-dong 800, Seo-gu, Daejeon 302-729, Republic of Korea
3. Department of Bioresource Science, College of Agriculture, Ibaraki University; Ami-machi, Ibaraki 300-0393, Japan