1. Division of Microbial and Molecular Ecology, Institute of Life Sciences and the Moshe Shilo Center for Marine Biogeochemistry, The Hebrew University of Jerusalem, Jerusalem 91904, Israel
2. Department of Microbiology and Parasitology, Faculty of Pharmacy, University of Seville, Seville 41012, Spain
3. Noda Institute for Scientific Research, 399 Noda, Noda-shi, Chiba-ken 278-0037, Japan