1. Estación Biológica de Doñana, Consejo Superior de Investigaciones Científicas (CSIC), Avenida de Américo Vespucio s/n, 41092 Sevilla, Spain
2. Department of Biology, University of Western Ontario, N6A 5B7, London, Ontario, Canada
3. School of Life Sciences, University of KwaZulu-Natal, P/Bag X01, Scottsville, Pietermaritzburg 3209, South Africa