1. 1United States Department of Agriculture, Agricultural Research Service, Arthropod-borne Animal Diseases Research Unit, 1515 College Avenue, Manhattan, KS 66502, USA
2. 2Southeastern Cooperative Wildlife Disease Study, Department of Population Health, College of Veterinary Medicine, University of Georgia, 501 D.W. Brooks Drive, Athens, GA 30602, USA
3. 3Department of Veterinary and Biomedical Sciences, South Dakota State University, SAR 125, Box 2175, Brookings, SD 57007, USA
4. 4United States Department of Agriculture, Animal-Plant Health Inspection Service, National Veterinary Service Laboratories, Diagnostic Virology Laboratory, PO Box 844, Ames, IA 50010, USA