1. School of Biological Sciences, Royal Holloway, University of London, Egham, Surrey TW20 OEX, UK
2. Department of Bacteriology, Animal Health and Veterinary Laboratories Agency, Addlestone, Surrey KT15 3NB, UK
3. Department of Food and Nutritional Sciences, University of Reading, Reading, Berkshire RG6 6AP, UK
4. Pathology Unit, Animal Health and Veterinary Laboratories Agency, Addlestone, Surrey KT15 3NB, UK
5. Faculty of Health and Medical Sciences, University of Surrey, Guilford, Surrey GU2 7XH, UK