1. Department of Applied Biology and Chemistry, Faculty of Applied Bioscience, Tokyo University of Agriculture, Sakuragaoka 1-1-1, Setagaya-ku, Tokyo 156-8502, Japan
2. R & D Department, Higeta Shoyu Co., Ltd, Choshi, Chiba 288-8680, Japan
3. Microbial Properties Research, National Center for Agricultural Utilization Research, US Department of Agriculture, Peoria, IL 61604, USA