1. Department of Food Science, Cornell University, Ithaca, NY 14853, USA
2. Department of Microbiology, Cornell University, Ithaca, NY 14853, USA
3. Present address: Department of Systems Biology, Columbia University, New York, NY 10032, USA
4. Present address: Department of Environmental and Forest Biology, SUNY College of Environmental Science and Forestry, Syracuse NY 13210, USA