1. Department of Immunobiology and BIO5 Institute, University of Arizona, Tucson, AZ 85719, USA
2. Department of Ecology and Evolutionary Biology, University of Arizona, Tucson, AZ 85721, USA
3. School of Animal and Comparative Biomedical Sciences, University of Arizona, Tucson, AZ 85721, USA
4. Department of Biology, Brooklyn College of the City University of New York, Brooklyn, NY 11210, USA
5. Department of Zoology, University of Oxford, Oxford OX1 3PS, UK