1. 1Department of Microbiology and Molecular Biology, College of Bioscience and Biotechnology, Chungnam National University, 99 Daehak-ro, Yuseong, Daejeon 34134, Republic of Korea
2. 2Bacterial Disease Division, Animal and Plant Quarantine Agency, 177 Hyeoksin 8-ro, Gimcheon 39660, Gyeongsangbuk-do, Republic of Korea
3. †Present address: Bacterial Disease Division, Animal and Plant Quarantine Agency, Gimcheon, Republic of Korea.
4. ‡Present address: Animal Disease Diagnostic Division, Animal and Plant Quarantine Agency, Gimcheon, Republic of Korea.