1. 1College of Life Sciences, Zhejiang Sci-Tech University, Hangzhou 310018, PR China
2. 2Key Laboratory of Marine Ecosystem and Biogeochemistry, Second Institute of Oceanography, State Oceanic Administration, Hangzhou 310012, PR China
3. 3College of Life Sciences, Zhejiang University, Hangzhou 310058, PR China
4. 4The Alexander Silberman Institute of Life Sciences, the Hebrew University of Jerusalem, The Edmond J. Safra Campus, 9190401 Jerusalem, Israel