1. 1College of Marine Life Sciences, Ocean University of China, Qingdao 266003, PR China
2. 2Sansha Track Ocean Coral Reef Conservation Research Institute, Sansha 573199, PR China
3. 3Laboratory for Marine Ecology and Environmental Science, Qingdao National Laboratory for Marine Science and Technology, Qingdao 266071, PR China
4. 4Key Laboratory of Marine Chemistry Theory and Technology, Ocean University of China, Ministry of Education, Qingdao 266100, PR China
5. 5College of Marine Geosciences, Ocean University of China, Qingdao 266100, PR China
6. 6Key Laboratory of Marine Genetics and Breeding (Ministry of Education), Ocean University of China, Qingdao 266003, PR China