1. State Key Laboratory of Microbial Technology, Marine Biotechnology Research Center, School of Life Science, Shandong University, Qingdao 266237, PR China
2. College of Marine Life Sciences, Institute for Advanced Ocean Study, Ocean University of China, Qingdao 266003, PR China
3. Laboratory for Marine Biology and Biotechnology, Qingdao National Laboratory for Marine Science and Technology, Qingdao 266235, PR China