1. Department of Microbiology and Molecular Biology, College of Bioscience and Biotechnology, Chungnam National University, 220 Gung-Dong, Yuseong, Daejeon 34134, Republic of Korea
2. Department of Biology, Inha University, Incheon 22212, Republic of Korea
3. Bacterial Resources Research Team, Freshwater Bioresources Research Division, Nakdonggang National Institute of Biological Resources, 137, Donam 2-Gil, Sangju, Gyeongsangbuk-Do 37242, Republic of Korea