1. School of Mathematics and Statistics, University of New South Wales, Sydney, New South Wales 2052, Australia;
2. Geophysical Fluid Dynamics Laboratory, National Oceanic and Atmospheric Administration, Princeton, New Jersey 08540, USA
3. Program in Atmospheric and Oceanic Sciences, Princeton University, Princeton, New Jersey 08540, USA
4. Laboratory of Ecology and Evolution of Plankton, Stazione Zoologica Anton Dohrn, 80121 Naples, Italy;
5. National Oceanography Centre, Southampton SO14 3ZH, United Kingdom