1. Department of Environmental Sciences, University of Virginia, Charlottesville, Virginia 22904, USA;
2. Ocean Acidification Program and Conservation Biology Division, Northwest Fisheries Science Center, National Marine Fisheries Service, National Oceanic and Atmospheric Administration, Seattle, Washington 98112, USA;
3. Ocean Conservancy, Washington, DC 20036, USA;
4. Department of Ecology and Evolutionary Biology, University of California, Santa Cruz, California 95064, USA;