1. Department of Biological and Environmental Science, University of Jyväskylä, Jyväskylä, Finland; e-mail: ossi.nokelainen@jyu.fi
2. CIBIO-InBIO Associate Laboratory, Research Center in Biodiversity and Genetic Resources, University of Porto, Vairão, Portugal; e-mail: boratyns@cibio.up.pt
3. School of Psychological Science, University of Bristol, Bristol, BS8 1TU, UK