1. Biodiversity Research Institute (IRBio), Universitat de Barcelona, E-08028 Barcelona, Catalonia, Spain email:
2. CIBIO-InBIO Associate Laboratory, Research Center in Biodiversity and Genetic Resources, University of Porto, PT-4485-661 Vairão, Portugal
3. Department of Zoology, University of Venda, Private Bag X5050, Thohoyandou, 0950, South Africa
4. Department of Comparative Anatomy, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30-387 Kraków, Poland