1. Department of Evolutionary Biology and Environmental Studies, University of Zurich, Winterthurerstrasse 190, CH-8057 Zürich, Switzerland.
2. Instituto Pirenaico de Ecología (IPE-CSIC), Apartado 13034, E-50080 Zaragoza, Spain.
3. Department of Life Sciences, University of Roehampton, Holybourne Avenue, London SW15 4JD, UK.