1. Department of Ecology, Evolution, and Environmental Biology, Columbia University, New York, NY, USA.
2. Department of Natural Resources and the Environment, Cornell University, Ithaca, NY, USA.
3. Deparment of Population Medicine and Diagnostic Sciences, Cornell University, Ithaca, NY, USA.
4. Department of Ecology and Evolutionary Biology, Cornell University, Ithaca, NY, USA.
5. USGS New York Cooperative Fish and Wildlife Unit, Department of Natural Resources, Cornell University, Ithaca, NY, USA.