1. Graduate School of Biological Sciences, Nara Institute of Science and Technology, Ikoma, Nara 630-0192, Japan.
2. RIKEN Center for Sustainable Resource Science, Yokohama, Kanagawa 230-0045, Japan.
3. National Institute for Basic Biology, Okazaki, Aich 444-8585, Japan.
4. School of Life Science, Graduate University for Advanced Studies, Okazaki, Aich 444-8585, Japan.