1. RIKEN Plant Science Center, Yokohama, Kanagawa 230–0045, Japan (M.Y., N.G., H.I., M.O., Y.N., N.N., M.K., T.D.); Graduate School of Biological Sciences, Nara Institute of Science and Technology, Ikoma, Nara 630–0192, Japan (M.Y., T.D.); RIKEN Biomass Engineering Program, Yokohama, Kanagawa 230–0045, Japan (M.O., T.D.); Graduate School of Bio-Applications and Systems Engineering, Tokyo University