1. Department of Integrative Biology, University of California, Berkeley, CA 94720–3140, USA.
2. Environmental Fluid Mechanics Laboratory, Department of Civil and Environmental Engineering, Stanford University, Stanford, CA 94305–4020, USA.
3. Department of Biological Sciences and the J. P. Scott Center for Neuroscience, Mind, and Behavior, Bowling Green State University, Bowling Green, OH 43403, USA.