1. Cancer Research UK Cambridge Research Institute (CRI), Li Ka Shing Centre, Robinson Way, Cambridge CB2 0RE, UK.
2. Department of Pathological Cell Biology, Medical Research Institute, Tokyo Medical and Dental University, 1-5-45 Yushima, Bunkyo-ku, Tokyo 113-8510, Japan.
3. Department of Oncology, University of Cambridge, Cambridge CB2 0RE, UK.
4. Life Sciences Institute, Department of Molecular and Integrative Physiology and Internal Medicine, University of Michigan, 210 Washtenaw Avenue, Ann Arbor, MI 48109, USA.