1. Department of Virus Research, John Innes Institute, Agricultural and Food Research Council (AFRC) Institute of Plant Science Research, Norwich 4R4 7UH, United Kingdom.
2. Department of Cell Biology, John Innes Institute, AFRC Institute of Plant Science Research, Norwich, NR4 7UH, United Kingdom.
3. Department of Biochemistry, University of Liverpool, Liverpool L69 3BX, United Kingdom.