1. Third Institute of Oceanography, Ministry of Natural Resources, Xiamen 361005, China.
2. Key Laboratory of Marine Ecological Conservation and Restoration, Ministry of Natural Resources, Xiamen 361005, China.
3. School of Biological Sciences and the Environment Institute, The University of Adelaide, Adelaide, Australia.
4. Hainan Academy of Ocean and Fisheries Sciences, Haikou 570125, China.