1. Department of Zoology, University of Otago, Post Office Box 56, Dunedin, New Zealand.
2. School of Biological Sciences, University of Bristol, Woodland Road, Bristol BS8 1UG, UK.
3. Environment Agency, Abu Dhabi, United Arab Emirates.
4. Institute of Natural Resources, Massey University, Private Bag 11222, Palmerston North, New Zealand.