1. Department of Chemistry, Biology and Marine Science, University of the Ryukyus, Nishihara, Okinawa 903-0123, Japan.
2. Centre for Coral Reef Biodiversity, James Cook University, Townsville, QLD 4811, Australia.
3. Department of Ecology, Evolution, and Marine Biology, University of California, Santa Barbara, CA 93106, USA.
4. Department of Biological Sciences, University of Delaware, Newark, DE 19716, USA.
5. Department of Paleontology, The Natural History Museum, Cromwell Road, London SW7 5BD, UK.