1. Department of Fisheries and Oceans, Institute of Ocean Sciences, 9860 West Saanich Road, Sidney, British Columbia V8L 4B2, Canada.
2. Research Institute for Global Change, Japan Agency for Marine-Earth Science and Technology, 2-15 Natsushima, Yokosuka, Kanagawa 237-0061, Japan.
3. Tokyo University of Marine Science and Technology, 4-5-7, Konan, Minato-ku, Tokyo 108-8477, Japan.