1. Institute of Marine Biology/Center of Excellence for the Oceans, National Taiwan Ocean University, Keelung 20224, Taiwan, Republic of China
2. School of Veterinary and Life Sciences, Murdoch University, Murdoch WA 6150, and Western Australian Herbarium, Science Division, Department of Parks and Wildlife, Locked Bag 104, Bentley Delivery Centre, WA 6983, Australia
3. Department of Botany, Smithsonian Institution, Washington, DC 20013-7012, USA