1. University of Girona, Faculty of Sciences, Department of Environmental Sciences, M. Aurèlia Capmany, 69, 17003 Girona, Spain
2. Phycology Research Group and Centre for Molecular Phylogenetics and Evolution, Ghent University, Ghent, Belgium
3. Department of Botany, National Museum of Nature and Science, 4-1-1 Amakubo, Tsukuba, Ibaraki 305-0005, Japan
4. Institute of Marine Biology, National Taiwan Ocean University, Keelung 20224, Taiwan, Republic of China