1. Department of Biological Sciences, Graduate School of Science, University of Tokyo, Hongo 7-3-1, Bunkyo-ku, Tokyo 113-0033, Japan
2. Applied Taxonomic Research Center, Department of Biology, Faculty of Science, Khon Kaen University, Nai-Muang, Muang District, Khon Kaen 40002, Thailand
3. Department of Biological Science and Chemistry, Faculty of Science, Graduate School of Medicine, Yamaguchi University, 1677-1 Yoshida, Yamaguchi 753-8512, Japan
4. Division of Biological Sciences, University of Montana, Missoula, MT 59812, USA
5. Center for Environmental Biology and Ecosystem Studies, National Institute for Environmental Studies, Onogawa 16-2, Tsukuba-shi, Ibaraki 305-8506, Japan