1. Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Hongo 7-3-1, Bunkyo-ku, Tokyo 113-0033, Japan
2. Department of Biological Sciences, Graduate School of Science, Kyoto University, Kitashirakawa-Oiwakecho, Sakyo-ku, Kyoto 606-8502, Japan
3. Center for Environmental Biology and Ecosystem Studies, National Institute for Environmental Studies, Onogawa 16-2, Tsukuba-shi, Ibaraki 305-8506, Japan
4. Department of Biological Science and Chemistry, Faculty of Science, Graduate School of Medicine, Yamaguchi University, 1677-1 Yoshida, Yamaguchi 753-8512, Japan
5. Applied Taxonomic Research Center, Department of Biology, Faculty of Science, Khon Kaen University, Nai-Muang, Muang District, Khon Kaen 40002, Thailand