1. Kyushu University Amakusa Marine Biological Laboratory, Tomioka 2231, Amakusa-Reihoku 863-2507, Japan
2. Kobe University Research Center for Inland Seas, Rokkodai, Kobe 657-8501, Japan
3. Takehara Fisheries Research Station, Setouchi Field Science Center, Hiroshima University, Takehara, Hiroshima 725-0024, Japan