1. Kyushu University Amakusa Marine Biological Laboratory Reihoku‐Amakusa Kumamoto863‐2507 Japan
2. Kobe University Research Center for Inland Seas Rokkodai Kobe657‐8501 Japan
3. Takehara Fisheries Research Station Setouchi Field Science Center Hiroshima University Takehara Hiroshima725‐0024 Japan
4. National Research Institute of Fisheries and Environment of Inland Sea Fishery Research Agency Hatsukaichi Hiroshima739‐0452 Japan
5. Research Institute on Subtropical Ecosystems 252 Yaga Nago Okinawa905‐1631 Japan